Laurent Ballesta was selected as the winner of the competition following the submission of his “enigmatic image” Creation, which captures camouflage groupers exiting their milky cloud of eggs and sperm in Fakarava, French Polynesia. Take a look at all the other winners in various categories in our photo gallery.
13. Seaweed
Rich reflections by Justin Gilligan showing the reflection of a marine ranger among the seaweed in Australia, which won Wildlife Photographer of the Year: Plants and Fungi Award.
14. Ghost pipefish
Bedazzled by Alexander Mustard showing a ghost pipefish hiding among the arms of a feather star, which won Wildlife Photographer of the Year: Natural Artistry Award.
15. Cichlid fish
Two adult male cichlid fish fighting jaw to jaw over a snail shell in Lake Tanganyika, Africa, which won Wildlife Photographer of the Year: Portfolio Award. Angel Fitor/Wildlife Photographer of the Year/PA Media
16. Reindeer
Head to head by Stefano Unterthiner showing a two Svalbard reindeer battle for control of a harem, which won Wildlife Photographer of the Year: Behaviour: Mammals Award.